提出问题：However, other than F. vesca, which is universally accepted one of its diploid ancestors, its other early diploid progenitors remain unclear. 

主要研究：Here, we performed comparative analyses of the genomes of five diploid strawberries, F. iinumae, F. vesca, F. nilgerrensis, F. nubicola, and F. viridis, of which the latter three are newly sequenced.

结果1-比较基因组：We found that the genomes of these species share highly conserved gene content and gene order.

结果2-栽培种草莓的祖先：Using an alignment-based approach, we show that F. iinumae and F. vesca are the diploid progenitors to the octoploid F. × ananassa, whereas the other three diploids that we analyzed in this study are not parental species.

结果3-草莓属系统发育：We generated a fully resolved, dated phylogeny of Fragaria and determined that the genus arose ca. 6.37 million years ago.

结论：Our results effectively resolve conflicting hypotheses regarding the putative diploid progenitors of the cultivated strawberry, establish a reliable backbone phylogeny for the genus, and provide genetic resources for molecular breeding.