Long distance dispersal was advocated in the paper as in many other case studies.  I prefer this hypothesis to explain the biogeography of Solanaceae as well.  A summary is badly needed to clear how many taxa share the similar distribution pattern and if they have similar ages?  Many of them seems to formed from Eocene to Miocene.  Maybe the time of the close of the boreatropic flora need to be reexamined as well?