| National Institutes of Health, United States | 32986 | 475602 |
| Harvard University | 28507 | 375681 |
| Chinese Academy of Sciences | 81368 | 364745 |
| University of Cambridge | 28029 | 289248 |
| University of Tokyo | 44374 | 272947 |
| University of California Berkeley | 27451 | 266083 |
| Stanford University | 24828 | 254679 |
| Max Planck Gesellschaft | 34672 | 253277 |
| University of California San Diego | 25914 | 249507 |
| Kyoto University | 44059 | 248162 |
| University of California Los Angeles | 21272 | 222249 |
| Massachusetts Institute of Technology | 23198 | 216658 |
| University of Oxford | 26965 | 196943 |
| University of Washington | 17488 | 192960 |
| Osaka University | 36898 | 191493 |
| University of Toronto | 23300 | 184114 |
| Pennsylvania State University | 17434 | 181517 |
| University of Michigan | 22497 | 180643 |
| University of Wisconsin Madison | 19327 | 177107 |
| University of Minnesota | 21003 | 170187 |
| Yale University | 16612 | 162686 |
| Scripps Research Institute | 13621 | 159075 |
| Swiss Federal Institute of Technology Zurich | 19130 | 157811 |
| Tohoku University | 33276 | 155749 |
| University of California San Francisco | 10336 | 155385 |
| University of Pennsylvania | 15235 | 154883 |
| Imperial College London | 18703 | 154048 |
| Washington University in St. Louis | 11260 | 153369 |
| Cornell University | 15876 | 150728 |
| University of Illinois Urbana Champaign | 18253 | 144582 |
| Columbia University | 13849 | 144360 |
| ERNET India | 38558 | 136633 |
| University of North Carolina Chapel Hill | 13461 | 134493 |
| University of Georgia | 9486 | 131417 |
| Hokkaido University | 25593 | 130879 |
| University of Florida | 17734 | 128427 |
| University of California Davis | 15888 | 128169 |
| Johns Hopkins University | 10786 | 124628 |
| University College London | 13959 | 123975 |
| Kyushu University | 23252 | 122967 |
| Nagoya University | 20733 | 121134 |
| Northwestern University | 15572 | 120049 |
| National Institute of Advanced Industrial Science & Technology, Japan | 23243 | 120040 |
| University of Manchester | 18138 | 118276 |
| Universität Basel | 14273 | 117806 |
| University of Texas Austin | 16669 | 115942 |
| Consejo Superior de Investigaciones Científicas | 23261 | 115469 |
| University of Massachusetts | 16033 | 114472 |
| Karolinska Institute | 11129 | 112678 |
| Russian Academy of Sciences | 74886 | 111777 |
| Ohio State University | 17829 | 109490 |
| California Institute of Technology | 12483 | 109403 |
| Duke University | 10620 | 108581 |
| Tokyo Institute of Technology | 25953 | 107348 |
| Technical University Munich | 13832 | 106925 |
| University of Pittsburgh | 12015 | 105101 |
| Purdue University | 16563 | 103458 |
| National University of Singapore | 13608 | 100654 |
| University of Alberta | 13987 | 99476 |
| McGill University | 13807 | 99245 |
| Centre National de la Recherche Scientifique | 13817 | 99055 |
| Case Western Reserve University | 11002 | 98970 |
| Ludwig Maximilian University of Munich | 13544 | 98802 |
| University of British Columbia | 13401 | 98487 |
| Zhejiang University | 21341 | 98262 |
| European Molecular Biology Laboratory | 2482 | 97947 |
| Texas A&M University | 17909 | 96074 |
| Agricultural Research Service | 11791 | 95937 |
| University of Southern California | 10919 | 95301 |
| Uppsala University | 11369 | 94355 |
| University of Utah | 12367 | 93654 |
| University of Bristol | 11259 | 93219 |
| University of California Irvine | 10495 | 93102 |
| Consiglio Nazionale delle Ricerche | 16978 | 92859 |
| University of Copenhagen | 11376 | 91481 |
| Institut National de la Recherche Agronomique INRA | 10554 | 89473 |
| Seoul National University | 15061 | 88646 |
| Atomic Energy Commission | 14297 | 87837 |
| Utrecht University | 11998 | 86772 |
| Lund University | 12136 | 86562 |
| Georgia Institute of Technology | 11401 | 86116 |
| University of Maryland | 9947 | 85351 |
| Tsinghua University Beijing, China | 16938 | 84893 |
| Vanderbilt University | 9967 | 84784 |
| Stockholm University | 9569 | 84559 |
| University of Groningen | 8952 | 83421 |
| Michigan State University | 10303 | 82816 |
| University of California Santa Barbara | 9888 | 82235 |
| University of Chicago | 9029 | 80029 |
| Heidelberg University | 10052 | 79852 |
| Rutgers, The State University of New Jersey | 10225 | 79458 |
| University of Sydney | 10764 | 77525 |
| Princeton University | 8963 | 76875 |
| University of Leeds | 12388 | 76004 |
| Catholic University of Leuven | 12580 | 75498 |
| University of Queensland | 9247 | 75498 |
| University of Colorado Boulder | 9315 | 74546 |
| Emory University | 8132 | 74101 |
| University of Edinburgh | 9875 | 72940 |
| Peking University | 15367 | 72718 |
