| 1 | Harvard University, United States of America (USA) | 610.22 | 612.01 | 1488 | 2.2% |
| 2 | National Institutes of Health (NIH), United States of America (USA) | 314.47 | 341.92 | 828 | 10.8% |
| 3 | Max Planck Society, Germany | 281.15 | 272.31 | 859 | -1.3% |
| 4 | Stanford University, United States of America (USA) | 236.22 | 271.97 | 631 | 17.3% |
| 5 | Chinese Academy of Sciences (CAS), China | 212.66 | 206.77 | 667 | -0.9% |
| 6 | Massachusetts Institute of Technology (MIT), United States of America (USA) | 208.5 | 200.16 | 740 | -2.2% |
| 7 | Yale University, United States of America (USA) | 193.05 | 175.03 | 402 | -7.6% |
| 8 | University of California, San Diego (UC San Diego), United States of America (USA) | 165.91 | 172.73 | 480 | 6.1% |
| 9 | University of California San Francisco (UCSF), United States of America (USA) | 193.93 | 172.07 | 491 | -9.6% |
| 10 | University of Pennsylvania (Penn), United States of America (USA) | 143.62 | 169.26 | 436 | 20.1% |
| 11 | Columbia University in the City of New York (CU), United States of America (USA) | 157.43 | 168.36 | 412 | 9.0% |
| 12 | French National Centre for Scientific Research (CNRS), France | 176.06 | 167.09 | 1096 | -3.3% |
| 13 | University of Oxford, United Kingdom (UK) | 193.18 | 165.79 | 503 | -12.6% |
| 14 | Johns Hopkins University (JHU), United States of America (USA) | 152.71 | 152.75 | 420 | 1.9% |
| 15 | Washington University in St. Louis (WUSTL), United States of America (USA) | 143.41 | 151.06 | 325 | 7.3% |
| 16 | University of Cambridge, United Kingdom (UK) | 168.34 | 149.93 | 482 | -9.3% |
| 17 | University of Michigan (U-M), United States of America (USA) | 122.44 | 147.51 | 365 | 22.7% |
| 18 | University of Toronto (U of T), Canada | 147.06 | 144.92 | 371 | 0.4% |
| 19 | The University of Texas Southwestern Medical Center (UT Southwestern Medical Center), United States of America (USA) | 113.71 | 139.62 | 267 | 25.1% |
| 20 | Cornell University, United States of America (USA) | 130.34 | 138.22 | 426 | 8.0% |
| 21 | University of California Berkeley (UC Berkeley), United States of America (USA) | 144.57 | 136.21 | 349 | -4.0% |
| 22 | University College London (UCL), United Kingdom (UK) | 130.03 | 124.85 | 498 | -2.2% |
| 23 | New York University (NYU), United States of America (USA) | 115.75 | 121.28 | 321 | 6.8% |
| 24 | University of California Los Angeles (UCLA), United States of America (USA) | 135.97 | 120.37 | 322 | -9.8% |
| 25 | University of Washington (UW), United States of America (USA) | 129.23 | 116.03 | 365 | -8.5% |
| 26 | Helmholtz Association of German Research Centres, Germany | 112.9 | 103.01 | 586 | -7.0% |
| 27 | Northwestern University (NU), United States of America (USA) | 91.71 | 102.14 | 251 | 13.5% |
| 28 | Memorial Sloan Kettering Cancer Center (MSKCC), United States of America (USA) | 84.47 | 102.05 | 258 | 23.1% |
| 29 | Duke University, United States of America (USA) | 139.64 | 101.24 | 317 | -26.1% |
| 30 | University of North Carolina at Chapel Hill (UNC), United States of America (USA) | 96.62 | 100.68 | 273 | 6.2% |
| 31 | The University of Chicago (UChicago), United States of America (USA) | 98.78 | 99.8 | 253 | 2.9% |
| 32 | University of Wisconsin-Madison (UW-Madison), United States of America (USA) | 100.45 | 99.57 | 202 | 1.0% |
| 33 | The Rockefeller University, United States of America (USA) | 91.3 | 96.37 | 232 | 7.5% |
| 34 | The University of Tokyo (UTokyo), Japan | 100.12 | 93.17 | 251 | -5.2% |
| 35 | Peking University (PKU), China | 58.95 | 88.11 | 225 | 52.3% |
| 36 | The University of Edinburgh, United Kingdom (UK) | 72.1 | 85.2 | 253 | 20.4% |
| 37 | Baylor College of Medicine (BCM), United States of America (USA) | 88.28 | 82.63 | 264 | -4.6% |
| 38 | Vanderbilt University (VU), United States of America (USA) | 82.31 | 82.17 | 220 | 1.7% |
| 39 | Princeton University, United States of America (USA) | 62.81 | 81.97 | 172 | 33.0% |
| 40 | National Institute for Health and Medical Research (INSERM), France | 79.64 | 77.15 | 707 | -1.3% |
| 41 | California Institute of Technology (Caltech), United States of America (USA) | 57.74 | 74.72 | 155 | 31.8% |
| 42 | The Scripps Research Institute (TSRI), United States of America (USA) | 81.47 | 74.67 | 208 | -6.6% |
| 43 | The University of Texas MD Anderson Cancer Center, United States of America (USA) | 87.8 | 74.55 | 241 | -13.5% |
| 44 | University of Pittsburgh (Pitt), United States of America (USA) | 94.63 | 74.4 | 205 | -19.9% |
| 45 | McGill University, Canada | 90.95 | 73.4 | 230 | -17.8% |
| 46 | University of Zurich (UZH), Switzerland | 86.37 | 73.1 | 251 | -13.8% |
| 47 | Weizmann Institute of Science (WIS), Israel | 79.75 | 72.86 | 145 | -6.9% |
| 48 | Imperial College London (ICL), United Kingdom (UK) | 90.68 | 72.71 | 397 | -18.3% |
| 49 | University of California Davis (UC Davis), United States of America (USA) | 64.98 | 71.11 | 212 | 11.5% |
| 50 | Ludwig Maximilians University of Munich (LMU), Germany | 54.86 | 70.58 | 264 | 31.1% |
| 51 | NIH National Cancer Institute (NCI), United States of America (USA) | 79.24 | 68.37 | 210 | -12.1% |
| 52 | Karolinska Institute (KI), Sweden | 67.92 | 67.98 | 295 | 2.0% |
| 53 | Medical Research Council (MRC), United Kingdom (UK) | 55.68 | 67.21 | 290 | 23.0% |
| 54 | Swiss Federal Institute of Technology Zurich (ETH Zurich), Switzerland | 70.21 | 66.61 | 205 | -3.3% |
| 55 | Emory University, United States of America (USA) | 67.5 | 65.35 | 182 | -1.4% |
| 56 | University of Copenhagen (UCPH), Denmark | 67.1 | 63.27 | 217 | -3.9% |
| 57 | Mount Sinai Health System (MSHS), United States of America (USA) | 70.51 | 63.02 | 248 | -8.9% |
| 58 | RIKEN, Japan | 54.36 | 60.04 | 198 | 12.5% |
| 59 | University of Minnesota (UMN), United States of America (USA) | 64.71 | 58.71 | 187 | -7.6% |
| 60 | The University of Queensland (UQ), Australia | 45.48 | 58.49 | 189 | 31.0% |
| 61 | Rutgers, The State University of New Jersey (RU), United States of America (USA) | 62.64 | 57.83 | 141 | -5.9% |
| 62 | University of Illinois at Urbana-Champaign (UIUC), United States of America (USA) | 41.89 | 57.61 | 135 | 40.1% |
| 63 | Heidelberg University (Uni Heidelberg), Germany | 49.97 | 57.54 | 214 | 17.3% |
| 64 | Shanghai Institutes for Biological Sciences (SIBS), CAS, China | 68.06 | 57 | 228 | -14.7% |
| 65 | Tsinghua University (TH), China | 50.93 | 56.68 | 182 | 13.4% |
| 66 | The University of Texas at Austin (UT Austin), United States of America (USA) | 65.1 | 55.87 | 127 | -12.6% |
| 67 | University of Southern California (USC), United States of America (USA) | 56.54 | 55.8 | 178 | 0.5% |
| 68 | King's College London (KCL), United Kingdom (UK) | 46.37 | 55.56 | 323 | 22.1% |
| 69 | Kyoto University, Japan | 53.68 | 55.33 | 157 | 5.0% |
| 70 | Spanish National Research Council (CSIC), Spain | 51.77 | 54.41 | 269 | 7.1% |
| 71 | University of Utah (Utah), United States of America (USA) | 44.57 | 52.26 | 147 | 19.4% |
| 72 | The Pennsylvania State University (Penn State), United States of America (USA) | 54.45 | 51.31 | 125 | -4.0% |
| 73 | University of Massachusetts Medical School (UMass Medical School), United States of America (USA) | 54.84 | 50.83 | 159 | -5.6% |
| 74 | The Ohio State University (OSU), United States of America (USA) | 59.61 | 50.35 | 137 | -13.9% |
| 75 | University of Basel (UB), Switzerland | 48.67 | 50.25 | 172 | 5.2% |
| 76 | National University of Singapore (NUS), Singapore | 39 | 49.55 | 197 | 29.4% |
| 77 | MRC Laboratory of Molecular Biology, United Kingdom (UK) | 40.7 | 48.39 | 90 | 21.1% |
| 78 | Fudan University, China | 32.84 | 47.52 | 145 | 47.4% |
| 79 | Hebrew University of Jerusalem (HUJI), Israel | 44.18 | 47.18 | 111 | 8.8% |
| 80 | Albert Einstein College of Medicine (Einstein), United States of America (USA) | 47.93 | 47.14 | 143 | 0.2% |
| 81 | University of Colorado Denver | Anschutz Medical Campus (CU Anschutz), United States of America (USA) | 46.58 | 46.99 | 127 | 2.8% |
| 82 | University of California Irvine (UCI), United States of America (USA) | 52.43 | 46.77 | 117 | -9.1% |
| 83 | The University of British Columbia (UBC), Canada | 58.18 | 46.54 | 157 | -18.5% |
| 84 | Indiana University (IU), United States of America (USA) | 50.58 | 45.98 | 137 | -7.4% |
| 85 | Zhejiang University (ZJU), China | 42.89 | 45.46 | 145 | 8.0% |
| 86 | University of Freiburg (Uni Freiburg), Germany | 34.19 | 43.88 | 148 | 30.8% |
| 87 | Boston University (BU), United States of America (USA) | 50.61 | 43.83 | 191 | -11.8% |
| 88 | University of Cincinnati (UC), United States of America (USA) | 43.88 | 43.67 | 118 | 1.4% |
| 89 | Shanghai Jiao Tong University (SJTU), China | 30.74 | 43.14 | 155 | 43.0% |
| 90 | University of Montreal (UdeM), Canada | 44.08 | 43.05 | 139 | -0.5% |
| 91 | Oregon Health and Science University (OHSU), United States of America (USA) | 50.89 | 42.72 | 113 | -14.5% |
| 92 | Leibniz Association, Germany | 46.75 | 42.69 | 272 | -7.0% |
| 93 | Osaka University, Japan | 61.16 | 42.3 | 125 | -29.5% |
| 94 | Utrecht University (UU), Netherlands | 41.3 | 42.25 | 179 | 4.2% |
| 95 | Istituto Di Ricovero e Cura a Carattere Scientifico (IRCCS), Italy | 41.63 | 42.07 | 201 | 3.0% |
| 96 | Texas A&M University (TAMU), United States of America (USA) | 32.12 | 41.67 | 107 | 32.2% |
| 97 | European Molecular Biology Laboratory (EMBL), Germany | 42.39 | 41.29 | 208 | -0.7% |
| 98 | Seoul National University (SNU), South Korea | 37.23 | 40.88 | 101 | 11.9% |
| 99 | Case Western Reserve University (CWRU), United States of America (USA) | 53.21 | 40.04 | 117 | -23.3% |
| 100 | Catholic University of Leuven (KU Leuven), Belgium | 42.49 | 39.95 | 141 | -4.2%
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