提出问题：TERMINAL FLOWER 1 (TFL1) and FLOWERING LOCUS T (FT) and their orthologs antagonistically regulate these life history traits, yet their mechanism of action, antagonism and targets remain poorly understood.

结果1-TFL1被FD招募：Here, we show that TFL1 is recruited to thousands of loci by the bZIP transcription factor FD.

结果2-TFL1与FT共同的靶基因LFY：We identify the master regulator of floral fate, LEAFY (LFY) as a target under dual opposite regulation by TFL1 and FT and uncover a pivotal role of FT in promoting flower fate via LFY upregulation.

结果3-FT与TFL1拮抗基础：We provide evidence that the antagonism between FT and TFL1 relies on competition for chromatin-bound FD at shared target loci.

结果4-TFL1-FD复合物：Direct TFL1-FD regulated target genes identify this complex as a hub for repressing both master regulators of reproductive development and endogenous signalling pathways.

结论：Our data provide mechanistic insight into how TFL1-FD sculpt inflorescence architecture, a trait important for reproductive success, plant architecture and yield.